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Most crustaceans are di or tri chromatic, but some species can detect many more colors. For example, stomatopods have up to 16 different visual pigments and Daphnia, a branchiopod, has 46 opsins (the primary component of visual pigments) (Porter, 2009; Colbourne, 2011). Interestingly, Daphnia has only 22 ommatidia and is only known to have 4color vision, plus the ability to detect polarized light. Microarray data suggest Structure, Development and Evolution of Crustacean Eyes 21 that at least some of the more similar opsins are expressed in overlapping but unique patterns over the course of the animal’s life (Colbourne, 2011).

And Marshall, J. (2012). Understanding the complex visual system of Mantis Shrimps (Stomatopods); a new form of colour processing? , and Kojima, T. (2000). Requirements of Lim1, a Drosophila LIM-homeobox gene, for normal leg and antennal development. Development 127, 4315–4323. 36 Ajna S. , and Brendonck, L. (2012). Toward a Global Phylogeny of the “Living Fossil” Crustacean Order of the Notostraca. PLoS ONE 7, e34998. Vogt, K. (1975). Zur Optik des Flusskrebsauges. Z. Naturforsch. C 30, 691.

And Hofbauer, A. (2002). The Extraretinal Eyelet of Drosophila: Development, Ultrastructure, and Putative Circadian Function. J. Neurosci. 22, 9255–9266. , and Nusse, R. (1993). Cell patterning in the Drosophila segment: engrailed and wingless antigen distributions in segment polarity mutant embryos. Dev. Suppl. 105–114. L. (1990). The Ultrastructure of the Compound Eye of Two Species of Marine Ostracodes (Crustacea: Ostracoda: Cypridinidae). Acta Zoologica 71, 217–223. L. (2004). Molecular and Comparative Genetics of Mental Retardation.

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